Image: Nate Hepler

Nate Hepler

Technical Advisor


Nathan Hepler, Ph.D. is a technical advisor in the Litigation practice of Paul Hastings and is based in the firm’s New York office.  Dr. Hepler earned his Bachelor of Science degree in Biochemistry and Molecular Biology from Lebanon Valley College in 2013 and completed his Ph.D. in Plant Biology at The Pennsylvania State University in 2019.  Prior to entering graduate school, he completed a post-baccalaureate appointment at the National Institutes of Health.  He joins Paul Hastings following a postdoctoral appointment at The Pennsylvania State University.

Dr. Hepler’s research experience includes a broad range of biochemistry, molecular biology, microbiology, bioinformatics, and genetics.  He has co-authored multiple scientific publications, grants, and fellowships.  Dr. Hepler has also presented his research at several national conferences.



  • The Pennsylvania State University, Ph.D., 2019
  • Lebanon Valley College, B.S., 2013

Engagement & Publications

  • Hepler NK, Bowman A, Carey RE and Cosgrove DJ. 2020. Expansin loss is a common occurrence during adaptation to an aquatic environment. The Plant Journal. 101(3): 666-680
  • Hepler NK and Cosgrove DJ. 2019. Directed in vitro evolution of bacterial expansin BsEXLX1 for higher cellulose binding and its consequences for plant cell wall-loosening activities. FEBS Letters. 592(18): 2545-2555.
  • Pomerantsev AP, McCall RM, Chahoud M, Hepler NK, Fattah R and Leppla SH. 2017. Genome engineering in Bacillus anthracis using tyrosine site-specific recombinases. PLoS One. 22(8):e0183346.
  • Cosgrove DJ, Hepler NK, Wagner ER and Durachko DM. 2017. Measuring the Biomechanical Loosening Action of Bacterial Expansins on Paper and Plant Cell Walls. Methods Mol Biol. 1588:157-165.
  • Carey RE, Hepler NK and Cosgrove DJ. 2013. Selaginella moellendorffii has a reduced and highly conserved expansin superfamily with genes more closely related to angiosperms than to bryophytes. BMC Plant Biol. 13:4.

Practice Areas





The Pennsylvania State University, Ph.D. 2019

Lebanon Valley College, B.S. 2013

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